/* === Base === */
*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
  --cream: #FDF8F0;
  --dark: #1a1a1a;
  --dark-bg: #0f0f0f;
  --nautilus: #D4A574;
  --nautilus-light: #F5F0E8;
  --lobster: #8B2500;
  --lobster-light: #f0d0c0;
  --nervous: #F4D03F;
  --respiratory: #5DADE2;
  --digestive: #58D68D;
  --immune: #AF7AC5;
  --circulatory: #E74C3C;
  --skeletal: #FDEBD0;
  --endocrine: #F0B27A;
  --muscular: #F1948A;
  --hgt: #48C9B0;
  --sensory: #F5B041;
  --mnemonic: #85C1E9;
  --from-cc: #D4A574;
  --from-oc: #c0392b;
  --from-both: #8e44ad;
}

html { scroll-behavior: smooth; }
body {
  font-family: 'Inter', sans-serif;
  color: var(--dark);
  background: var(--cream);
  line-height: 1.6;
  font-weight: 300;
}

h1, h2, h3 { font-family: 'Cormorant Garamond', serif; font-weight: 600; }
em { font-family: 'Cormorant Garamond', serif; }

.container { max-width: 1100px; margin: 0 auto; padding: 0 24px; }
.container-wide { max-width: 1400px; margin: 0 auto; padding: 0 24px; }

.section { padding: 80px 0; }
.section-alt { background: #f5f0e8; }
.section-dark { background: var(--dark-bg); color: #e0e0e0; }
.section-dark h2 { color: #fff; }
.section-dark .section-intro { color: #aaa; }

.section-intro {
  font-size: 1.1rem;
  color: #666;
  max-width: 700px;
  margin: 0 auto 48px;
  text-align: center;
}

h2 {
  font-size: 2.4rem;
  text-align: center;
  margin-bottom: 12px;
}

/* === Language Toggle === */
.lang-toggle {
  position: fixed;
  top: 16px;
  right: 16px;
  z-index: 100;
  display: flex;
  gap: 4px;
  background: rgba(0,0,0,0.6);
  backdrop-filter: blur(8px);
  padding: 4px;
  border-radius: 8px;
}
.lang-btn {
  padding: 6px 14px;
  border: none;
  background: transparent;
  color: #888;
  font-size: 0.8rem;
  font-weight: 600;
  cursor: pointer;
  border-radius: 6px;
  transition: all 0.2s;
}
.lang-btn.active {
  background: var(--nautilus);
  color: #0a0e17;
}
.lang-btn:hover:not(.active) { color: #fff; }

/* === Hero === */
.hero {
  min-height: 100vh;
  display: flex;
  align-items: center;
  justify-content: center;
  text-align: center;
  background: #0a0e17;
  color: #fff;
  padding: 40px 24px;
  position: relative;
  overflow: hidden;
}
.hero-video {
  position: absolute;
  top: 50%; left: 50%;
  min-width: 100%; min-height: 100%;
  transform: translate(-50%, -50%);
  object-fit: cover;
  opacity: 0.35;
  z-index: 0;
}
.hero-overlay {
  position: absolute;
  inset: 0;
  background: linear-gradient(180deg, rgba(10,14,23,0.6) 0%, rgba(15,26,46,0.8) 100%);
  z-index: 1;
}
.hero-content { position: relative; z-index: 2; }

.hero-label {
  font-size: 0.85rem;
  letter-spacing: 3px;
  text-transform: uppercase;
  color: var(--nautilus);
  margin-bottom: 16px;
}

.hero h1 {
  font-size: clamp(2.4rem, 6vw, 4.2rem);
  line-height: 1.15;
  margin-bottom: 20px;
}

.hero-subtitle {
  font-size: 1.3rem;
  color: #aaa;
  margin-bottom: 12px;
}

.hero-desc {
  font-size: 1.05rem;
  color: #888;
  max-width: 550px;
  margin: 0 auto 40px;
}

.hero-species {
  display: flex;
  align-items: center;
  justify-content: center;
  gap: 32px;
  flex-wrap: wrap;
}

.species-badge {
  display: flex;
  align-items: center;
  gap: 12px;
  padding: 16px 24px;
  border-radius: 12px;
  font-size: 1rem;
  line-height: 1.3;
}
.species-badge small { font-size: 0.8rem; color: #aaa; }
.species-icon { font-size: 2.4rem; }

.nautilus { background: rgba(212,165,116,0.15); border: 1px solid rgba(212,165,116,0.3); }
.lobster { background: rgba(139,37,0,0.2); border: 1px solid rgba(139,37,0,0.4); }

.vs {
  font-family: 'Cormorant Garamond', serif;
  font-size: 1.6rem;
  color: #555;
  font-style: italic;
}

.hero-cta {
  display: inline-block;
  margin-top: 32px;
  padding: 12px 32px;
  border: 1px solid var(--nautilus);
  color: var(--nautilus);
  text-decoration: none;
  border-radius: 30px;
  font-size: 0.9rem;
  letter-spacing: 1px;
  transition: all 0.3s;
}
.hero-cta:hover {
  background: var(--nautilus);
  color: #0a0e17;
}

/* === Anatomy Image === */
.anatomy-image img {
  width: 100%;
  border-radius: 12px;
  box-shadow: 0 8px 40px rgba(0,0,0,0.15);
}

/* === Phylogeny Image === */
.phylo-image {
  max-width: 700px;
  margin: 0 auto;
}
.phylo-image img {
  width: 100%;
  border-radius: 12px;
  box-shadow: 0 8px 40px rgba(0,0,0,0.1);
}

/* === Panorama === */
.panorama-image {
  overflow-x: auto;
  -webkit-overflow-scrolling: touch;
}
.panorama-image img {
  width: 100%;
  min-width: 900px;
  border-radius: 12px;
  box-shadow: 0 8px 40px rgba(0,0,0,0.1);
}

/* === Video Player === */
.video-player {
  max-width: 800px;
  margin: 0 auto 40px;
}
.video-player video {
  width: 100%;
  border-radius: 12px;
  box-shadow: 0 8px 40px rgba(0,0,0,0.3);
}
.video-caption {
  text-align: center;
  font-size: 0.85rem;
  color: #888;
  margin-top: 12px;
  font-style: italic;
}

/* === Organ Gallery === */
.organ-gallery {
  display: grid;
  grid-template-columns: 1fr;
  gap: 24px;
}
.organ-gallery-item {
  border-radius: 12px;
  overflow: hidden;
  border: 1px solid #333;
  background: #1a1a1a;
  cursor: pointer;
  transition: transform 0.2s;
}
.organ-gallery-item:hover { transform: translateY(-2px); }
.organ-gallery-item img {
  width: 100%;
  display: block;
}
.organ-gallery-info {
  padding: 16px 20px;
  display: flex;
  justify-content: space-between;
  align-items: center;
}
.organ-gallery-info .organ-title {
  font-family: 'Cormorant Garamond', serif;
  font-size: 1.1rem;
  font-weight: 600;
  color: #fff;
}
.organ-gallery-info .organ-sub {
  font-size: 0.8rem;
  color: #888;
}
.organ-gallery-detail {
  display: none;
  padding: 0 20px 20px;
}
.organ-gallery-detail.open { display: block; }
.organ-gallery-detail .detail-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 16px;
}
@media (max-width: 600px) { .organ-gallery-detail .detail-grid { grid-template-columns: 1fr; } }
.organ-gallery-detail h4 {
  font-size: 0.85rem;
  margin-bottom: 6px;
  font-family: 'Cormorant Garamond', serif;
}
.organ-gallery-detail p {
  font-size: 0.8rem;
  color: #bbb;
  line-height: 1.5;
}
.organ-gallery-detail .file {
  font-size: 0.7rem;
  color: #666;
  font-family: monospace;
  margin-top: 4px;
}

/* === Taxonomy === */
.taxonomy-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 32px;
}
@media (max-width: 700px) { .taxonomy-grid { grid-template-columns: 1fr; } }

.taxonomy-card {
  padding: 32px;
  border-radius: 12px;
  border: 1px solid #ddd;
}
.nautilus-card { background: var(--nautilus-light); border-color: var(--nautilus); }
.lobster-card { background: var(--lobster-light); border-color: var(--lobster); }

.taxonomy-card h3 {
  font-size: 1.4rem;
  margin-bottom: 16px;
}

.taxonomy-card table {
  width: 100%;
  margin-bottom: 16px;
  border-collapse: collapse;
}
.taxonomy-card td {
  padding: 6px 0;
  border-bottom: 1px solid rgba(0,0,0,0.08);
}
.taxonomy-card td:first-child {
  font-size: 0.85rem;
  color: #888;
  text-transform: uppercase;
  letter-spacing: 1px;
  width: 100px;
}

.traits {
  list-style: none;
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
}
.traits li {
  font-size: 0.8rem;
  padding: 4px 10px;
  background: rgba(0,0,0,0.06);
  border-radius: 20px;
}

/* === Organ Systems === */
.organ-grid {
  display: grid;
  grid-template-columns: 1fr;
  gap: 20px;
}

.organ-card {
  display: grid;
  grid-template-columns: 200px 1fr 1fr;
  gap: 0;
  border-radius: 12px;
  overflow: hidden;
  border: 1px solid #333;
  background: #1a1a1a;
}
@media (max-width: 800px) {
  .organ-card { grid-template-columns: 1fr; }
}

.organ-label {
  padding: 20px;
  display: flex;
  flex-direction: column;
  justify-content: center;
  color: #fff;
  font-weight: 500;
}
.organ-label .organ-name {
  font-family: 'Cormorant Garamond', serif;
  font-size: 1.2rem;
  font-weight: 600;
  margin-bottom: 4px;
}
.organ-label .organ-bio {
  font-size: 0.75rem;
  color: #aaa;
}
.organ-label .organ-agent {
  font-size: 0.75rem;
  color: #888;
  margin-top: 2px;
}

.organ-species {
  padding: 20px;
  border-left: 1px solid #333;
}
.organ-species h4 {
  font-size: 0.85rem;
  margin-bottom: 8px;
  font-family: 'Cormorant Garamond', serif;
}
.organ-species p {
  font-size: 0.8rem;
  color: #bbb;
  line-height: 1.5;
}
.organ-species .file {
  font-size: 0.7rem;
  color: #666;
  font-family: monospace;
  margin-top: 6px;
}

/* === Phylogeny === */
.phylo-tree {
  max-width: 700px;
  margin: 0 auto;
  position: relative;
  padding-left: 80px;
}
.phylo-tree::before {
  content: '';
  position: absolute;
  left: 55px;
  top: 0;
  bottom: 0;
  width: 3px;
  background: linear-gradient(180deg, #ddd, var(--nautilus), var(--lobster));
  border-radius: 2px;
}

.phylo-era {
  position: relative;
  margin-bottom: 40px;
}

.phylo-year {
  position: absolute;
  left: -80px;
  top: 0;
  font-family: 'Cormorant Garamond', serif;
  font-size: 1.4rem;
  font-weight: 700;
  color: var(--nautilus);
  width: 55px;
  text-align: right;
}

.phylo-events { padding-left: 24px; }

.phylo-node {
  font-weight: 500;
  font-size: 0.95rem;
  margin-bottom: 6px;
  padding: 4px 0;
}
.phylo-node.epoch { color: #333; font-style: italic; }
.phylo-node.env-change {
  color: var(--digestive);
  font-size: 0.85rem;
  margin-top: 8px;
}
.phylo-node.convergence {
  color: var(--immune);
  font-weight: 600;
}

.phylo-species {
  font-size: 0.85rem;
  color: #888;
  margin-bottom: 4px;
}

.phylo-branches {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 12px;
  margin-top: 8px;
}

.phylo-branch {
  font-size: 0.82rem;
  padding: 12px;
  border-radius: 8px;
  line-height: 1.4;
}
.branch-a { background: rgba(212,165,116,0.12); border: 1px solid rgba(212,165,116,0.3); }
.branch-b { background: rgba(139,37,0,0.1); border: 1px solid rgba(139,37,0,0.25); color: var(--lobster); }

/* === Physiology === */
.physiology-grid {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 24px;
}
@media (max-width: 700px) { .physiology-grid { grid-template-columns: 1fr; } }

.physio-card {
  background: #1a1a1a;
  border-radius: 12px;
  padding: 24px;
  border: 1px solid #333;
}
.physio-card h3 {
  font-size: 1.2rem;
  color: #fff;
  margin-bottom: 16px;
}

.physio-table {
  width: 100%;
  border-collapse: collapse;
  font-size: 0.82rem;
}
.physio-table th {
  text-align: left;
  padding: 8px 6px;
  border-bottom: 1px solid #444;
  color: #aaa;
  font-weight: 500;
}
.physio-table td {
  padding: 8px 6px;
  border-bottom: 1px solid #222;
  color: #ccc;
}
.physio-table .strong { color: var(--digestive); }
.physio-table .mid { color: var(--sensory); }
.physio-table .weak { color: var(--circulatory); }
.physio-table .na { color: #555; }

/* === Ludex === */
.ludex-blocks {
  max-width: 700px;
  margin: 0 auto;
  display: flex;
  flex-direction: column;
  gap: 12px;
}

.block-row {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 12px;
}
@media (max-width: 600px) { .block-row { grid-template-columns: 1fr; } }

.block {
  padding: 20px;
  border-radius: 10px;
  text-align: center;
  border: 2px solid;
  opacity: 0.85;
}
.block span { display: block; font-weight: 600; font-size: 1rem; margin-bottom: 4px; }
.block small { font-size: 0.75rem; color: #888; }

.from-cc { border-color: var(--from-cc); background: rgba(212,165,116,0.08); }
.from-oc { border-color: var(--from-oc); background: rgba(192,57,43,0.08); }
.from-both { border-color: var(--from-both); background: rgba(142,68,173,0.08); }

.ludex-note {
  text-align: center;
  margin-top: 24px;
  font-size: 0.9rem;
  color: #888;
}

/* === Paper === */
.paper-info {
  text-align: center;
  max-width: 600px;
  margin: 0 auto;
}
.paper-info p { margin-bottom: 8px; }
.paper-download {
  display: inline-block;
  margin-top: 20px;
  padding: 12px 32px;
  border: 1px solid var(--nautilus);
  color: var(--nautilus);
  text-decoration: none;
  border-radius: 30px;
  font-size: 0.9rem;
  letter-spacing: 1px;
  transition: all 0.3s;
}
.paper-download:hover {
  background: var(--nautilus);
  color: var(--dark-bg);
}

/* === Footer === */
.footer {
  text-align: center;
  padding: 40px 24px;
  background: var(--dark-bg);
  color: #666;
  font-size: 0.85rem;
  border-top: 1px solid #222;
}
.footer p { margin-bottom: 4px; }